Life Mammalia Rodentia Sciuridae

1400x1912 (~398Kb) USA: Montana, Mineral Co., Restarea on I-90 (47°25'9N 115°37'26W), 6.8.2012, Photo © Markku Savela	1600x971 (~494Kb) USA: Montana, Mineral Co., Restarea on I-90 (47°25'9N 115°37'26W), 6.8.2012, Photo © Markku Savela	900x1503 (~254Kb) USA: Montana, Mineral Co., Restarea on I-90 (47°25'9N 115°37'26W), 6.8.2012, Photo © Markku Savela
1800x1132 (~578Kb) USA: Montana, Mineral Co., Restarea on I-90 (47°25'9N 115°37'26W), 6.8.2012, Photo © Markku Savela

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

• Spermophilus adocetus (Merriam, 1903) WC.Mexico (Jalisco, Michoacan, N.Guerrero)

  • adocetus (Merriam, 1903); Proc. Biol. Soc. Washginton, 16: 79; TL: Mexico, Michoacan, La Salada, 40 mi. S. Uruapan
  • infernatus
• Spermophilus annulatus Audubon & Bachman, 1842 Mexico (Nayarit - N.Guerrero). See [maps]

  • Spermophilus annulatus Audubon & Bachman, 1842; J. Acad. Nat. Sci. Phila., (1) 8: 319; TL: Mexico, Colima, Manzanillo
• Spermophilus atricapillus Bryant, 1889 Baja California. See [maps]

  • Spermophilus atricapillus Bryant, 1889; Proc. Calif. Acad. Sci. (2) 2: 26; TL: Mexico, Baja California Sur, Commondu

  • Otospermophilus atricapillus; [CoL]
• Spermophilus beecheyi (Richardson, 1829) W.Washington - Baja California. See [maps] Californian Ground Squirrel

  • beecheyi (Richardson, 1829); Fauna Boreali-Americana, ..., 1: 170; TL: California, Monterey Co., Monterey

  • Otospermophilus beecheyi; [CoL] ; [iNaturalist]

  800x592 (~170Kb) La Jolla Beach, San Diego, California, October 1998, Photo © Ralf Schmode

  694x600 (~143Kb) La Jolla Beach, San Diego, California, October 1998, Photo © Ralf Schmode

• Spermophilus variegatus (Erxleben, 1777) Utah, S.Nevada - SW.Texas - Mexico (Puebla). See [maps]

  • variegatus (Erxleben, 1777); Syst. Regn. Anim.: 421; TL: Mexico, Valley of Mexico, near Mexico City

  • Otospermophilus variegatus; [CoL] ; [iNaturalist]

• Spermophilus alashanicus Büchner, 1888 SC.Mongolia, Alsahan, N.China (E.Nan Shan). See [maps]

  • Spermophilus alashanicus Büchner, 1888; Wiss. Res. Przewalski cent. Asien Zool. Th. 1 (Suageth.): 11; TL: N.China, S. Alashan Desert
  • dilutus; (Mongolian Altai)
  • obscurus
  • siccus; (Shansi)

  • Spermophilus alashanicus; [CoL] ; [iNaturalist]
  • Spermophilus dauricus alashanicus; [MPR], 83
• Spermophilus armatus Kennicott, 1863 Utah - Montana, SE.Idaho, W.Wyoming. See [maps]

  • Spermophilus armatus Kennicott, 1863; Proc. Acad. Nat. Sci. Phila., 15: 158; TL: Wyoming, Uinta Co., foothils of Uinta Mts., near Fort Bridger
• Spermophilus beldingi Merriam, 1888 E.Oregon, SW.Idaho, NE.California, N.Nevada, NW.Utah. See [maps]

  • Spermophilus beldingi Merriam, 1888; Ann. N.Y. Acad. Sci. 4: 317; TL: California, Placer Co., Donner
• Spermophilus brunneus (Howell, 1928) WC.Idaho

  • brunneus (Howell, 1928); Proc. Biol. Soc. Washington, 41: 211; TL: Idaho, Adams Co., New Meadows
• Spermophilus citellus (Linnaeus, 1766) CEU, SEU, Asia Minor, Caucasus, Palestine. See [maps] European Souslik

  • Mus citellus Linnaeus, 1766; Syst. Nat. (edn. 12) 1: 80; TL: Austria

  • Spermophilus citellus; [CoL] ; [iNaturalist]

  identification uncertain 1024x714 (~88Kb) NW.Kazakshtan, near Atyrau, 28.2.2004, Photo © A. Akin Köksal

  identification uncertain 1024x729 (~146Kb) NW.Kazakshtan, near Atyrau, 28.2.2004, Photo © A. Akin Köksal

  • S. c. citellus

    • citillus
    • gradojevici
    • istricus
    • karamani
    • laskarevi
    • Citellus citellus martinoi Peshev, 1955; (Rhodopen Mts, Bulgaria)
    • Citellus citellus balcanicus Markov, 1957; (Lukorsko, Sofia dist, Bulgaria)
    • Citellus citellus thracius Musaloglu, 1964; (SE. Slope of Muraltepe, near Yenibedir, Lüleburgaz, Kirklareli, European Turkey)

    • Spermophilus citellus citellus; [MPR], 83; [CoL]
    • Spermophilus citellus gradojevici; [CoL]
    • Spermophilus citellus istricus; [CoL]
    • Spermophilus citellus martinoi; [CoL]
• Spermophilus dauricus Brandt, 1843 Transbaikalia, Mongolia, N.China. See [maps] Daurian Souslik

  • Spermophilus dauricus Brandt, 1843; Bull. Phys. Math. Acad. Sci. St. Petersb. 2: 379; TL: Tarei-Nor, c. 250 miles E. of L. Baikal
  • mongolicus; (Hopei)
  • ramosus; (Manchuria)
  • umbratus; (Mongolia)
  • yamashinai

  • Spermophilus dauricus; [MPR], 83; [CoL] ; [iNaturalist]
• Spermophilus elegans Kennicott, 1863 NE.Nevada, SE.Oregon, S.Idaho, SW.Montana - C.Colorado, W.Nebraska. See [maps]

  • Spermophilus elegans Kennicott, 1863; Proc. Acad. Nat. Sci. Phila., 15: 158; TL: Wyoming, Uinta Co., Fort Bridger
  • aureus
  • nevadensis
• Spermophilus xanthoprymnus Bennett, 1835 Transcaucasia, Turkey, Syria, Palestine. See [maps]

  • Spermophilus xanthoprymnus Bennett, 1835; Proc. Zool. Soc. Lond. 1835: 90; TL: NE.Turkey, Erzerum
  • concolor Thomas
  • schmidti

  • Spermophilus citellus xanthoprymnus; [MPR], 83
  • Spermophilus xanthoprymnus; [CoL] ; [iNaturalist]
• Spermophilus musicus Ménétries, 1832 N.Caucasus. See [maps]

  • Spermophilus musicus Ménétries, 1832; Cat. Raisonne Mus. Peale Phila.: 21; TL: Uchkulan, "Caucasus", foot of Mt Elbruz
  • boehmi

  • Spermophilus musicus; [CoL] ; [iNaturalist]
  • Spermophilus pygmaeus musicus; [MPR], 83
• Spermophilus pygmaeus (Pallas, 1779) Crimea, Caucasus - Kazakhstan. See [maps] Little Souslik

  • Spermophilus pygmaeus; [CoL] ; [iNaturalist]
  • S. p. pygmaeus

    • Mus citellus var. pygmaeus Pallas, 1779; Nova Spec. Quad. Glir. Ord.: 122; TL: Kazakh S.S.R, Between Ural and Emba Rivers
    • arenicola Rall
    • atricapilla
    • binominata
    • brauneri; (Crimea)
    • ellermani
    • ?flavescens
    • herbicola
    • kazakstanicus
    • kalabuchovi
    • musogaricus
    • nikolskii
    • orlovi Ellerman
    • pallidus Orlov & Feniuk
    • planicola; (Caucasus)
    • ralli Heptner
    • satunini
    • septentrionalis

    • Spermophilus pygmaeus brauneri; [CoL]
    • Spermophilus pygmaeus herbicolus; [CoL]
    • Spermophilus pygmaeus mugosaricus; [CoL]
    • Spermophilus pygmaeus pygmaeus; [MPR], 83; [CoL]
    • Spermophilus ralli; [CoL]
• Spermophilus relictus (Kashkarov, 1923) -. See [maps] Tien Shan Souslik

  • Citellus musicus relictus Kashkarov, 1923; Trans. Turk. Sci. Sco. 1: 185; TL: Talassk. Alatau, Kara-Bura and Kumysh-Tagh Valleys

  • Spermophilus relictus; [MPR], 84; [CoL] ; [iNaturalist]
  • Spermophilus relictus Kashkarov, 1923;
  • S. r. relictus W.Tien Shan (between Tashkent and Issuk Kul)

    • Spermophilus relictus relictus; [MPR], #84
  • S. r. ralli (Kuznetzov, 1948) C.Tien Shan (E.Issyk Kul)

    • Citellus relictus ralli Kuznetsov, 1948; ; TL: Issyk Kul
• Spermophilus richardsonii (Sabine, 1822) N.Great Plains (S.Alberta, S.Saskatchewan, S.Manitoba, Montana, North Dakota, NE.South Dakota, W.Minnesota, NW.Iowa). See [maps]

  • richardsonii (Sabine, 1822); Trans. Linn. Soc. Lond. 13: 589; TL: Saskatchewan, Carleton House
• Spermophilus suslicus (Güldenstaedt, 1770) SW.Russia (from Volga and Don west to SE. Poland, NE.Romania). See [maps] Spotted Souslik

  • Mus suslica Güldenstaedt, 1770; Nova Comm. Acad. Sci. Petrop. 14: 389; TL: Voronej Steppes, near Voronezh, Russia
  • averini
  • guttatus
  • guttulatus
  • leucopictus
  • meridioccidentalis
  • odessana
  • ognevi; (Rumania)
  • volhyensis; (E. Poland)
  • Citellus suslicus boristhenicus Pusanov, 1958; ; TL: Between the rivers Dnestr and Bug, Ukraine

  • Spermophilus suslicus boristhenicus; [CoL]
  • Spermophilus suslicus guttatus; [CoL]
  • Spermophilus suslicus suslicus; [CoL]
  • Spermophilus suslicus; [MPR], 83; [CoL] ; [iNaturalist]
• Spermophilus townsendii Bachman, 1839 Great Basin (E.Washington, SE.Oregon, S.Idaho, E.California, Nevada, W.Utah). See [maps]

  • Spermophilus townsendii Bachman, 1839; J. Acad. Nat. Sci. Philadelphia, 8 (1): 61; TL: Washington, Wall Walla Co., near Wallula
  • ?mollis
  • ?vigilis
• Spermophilus washingtoni (Howell, 1938) SE.Washington, NE.Oregon. See [maps]

  • washingtoni (Howell, 1938); N. Am. Fauna 56: 69; TL: Washington, Walla Walla Co., Touchet

• Spermophilus columbianus (Ord, 1815) from (SE.British Columbia, W.Alberta) - to (NE.Oregon, Idaho, Montana). See [maps]

  • columbianus (Ord, 1815); in Guthrie, A New Geogr., Hist. Compl. Grammar, Phila, 2nd ed., 2: 292; TL: Idaho, Idaho Co., between the forks of the Clearwater and Kooskoskie Rivers

  • Urocitellus columbianus; [CoL] ; [iNaturalist]
• Spermophilus parryi (Richardson, 1825) NW.Canada, Alaska. See [maps] Arctic Souslik

  • Arctomys Parryii Richardson, 1825; in Parry, Journal of a second voyage for the discovery of a North-west Passage (Zool. App.): 316; TL: Hudson Bay, Candada

  • Spermophilus parryi; [MPR], 85
  • Spermophilus parryi Richardson, 1825;
  • S. p. leucostictus Brandt, 1844 NE.Siberia

    • Spermophilus parryi leucostictus Brandt, 1844; ; TL: Okhotsk River
    • buxtoni

    • Spermophilus parryi leucostictus; [MPR], 85
  • S. p. stejnegeri (Allen, 1903) Kamchatka

    • Spermophilus parryi stejnegeri; [MPR], 85
  • S. p. janensis (Ognev, 1937) River Jana

    • Spermophilus parryi janensis; [MPR], 85
  • S. p. coriakorum (Portenko, 1963) NE.Siberia (Koryak Mts.)

    • Citellus undulatus coriakorum Portenko, 1963; ; TL: Koryak Mts.

    • Spermophilus parryi coriakorum; [MPR], 85
• Spermophilus undulatus (Pallas, 1779) Tien Shan - Altai, Mongolia, Amur, Heilungkiang, Sinkiang. See [maps] Long-tailed Souslik

  • Mus citellus var. undulatum Pallas, 1779; Nova Spec. Quad. Glir. Ord.: 122; TL: Lake Baikal, Selenga R.
  • altaicus; (Altai)
  • eversmanni; (Altai)
  • intercedens; (E.Transbaikalia)
  • jacutensis; (Yakutsk)
  • menzbieri; (Upper Amur)
  • stramineus; (NW.Mongolia)
  • transbaicalikus

  • Spermophilus undulatus; [MPR], 84
  • Spermophilus undulatus Pallas, 1779;
  • Urocitellus undulatus; [CoL] ; [iNaturalist]

• Spermophilus erythrogenys Brandt, 1841 E.Kazakhstan, SW.Siberia, Sinkiang, Mongolia, Inner Mongolia. See [maps]

  • Spermophilus erythrogenys Brandt, 1841; Bull. Acad. Sci. St. Petersb.: 43; TL: Tomsk, foothills of Altai Mts., near Barnaul

  • Spermophilus erythrogenys; [CoL] ; [iNaturalist]
  • S. e. erythrogenys

    • Citellus eryhtrogenys brunnescens Belyaev, 1954

    • Spermophilus major erythrogenys; [MPR], 84
  • ?S. e. brevicauda Brandt, 1843

    • Spermophilus erythrogenys brevicauda Brandt, 1843; ; TL: SE.Kazakhstan
    • intermedius
    • carruthersi
    • pallicauda; (Mongolia,Inner Mongolia)
    • ?selevini
    • Citellus major heptneri Vasilieva, 1964; ; TL: ?Tien Shan
    • Citellus pygmaeus iliensis Belyaev, 1945; ; TL: River Ili

    • Spermophilus brevicauda; [CoL] ; [iNaturalist]
    • Spermophilus major brevicauda; [MPR], 84
• Spermophilus fulvus (Lichtenstein, 1823) Kazakhstan, Caspian Sea - Lake Balkash, NE.Iran, N.Afghanistan, Singkiang. See [maps] Large-toothed Souslik

  • Arctomys fulvus Lichtenstein, 1823; Naturh. Abh. Eversmann's Reise: 119; TL: E of Mugodzhary Mts, near Kuvandzhur River
  • concolor Fischer
  • concolor Geoffroy
  • giganteus
  • hypoleucos; (N.Iran)
  • nanus
  • nigrimontanus; (Karatau Mts.)
  • orlovi
  • oxianus; (S.Turkestan)
  • parthianus
  • Mus citellus maximus Pallas, 1779; Nova Spec. Quad. Glir. Ord.: 122; TL: Lower River Ural

  • Spermophilus fulvus fulvus; [CoL]
  • Spermophilus fulvus hypoleucos; [CoL]
  • Spermophilus fulvus oxianus; [CoL]
  • Spermophilus fulvus; [MPR], 85; [CoL] ; [iNaturalist]
• Spermophilus major (Pallas, 1779) N.Kasakstan (steppe Volga to Irtysh R.) Russet Souslik

  • Mus citellus var. major Pallas, 1779; ; TL: Samara, E of R. Volga

  • Spermophilus major; [MPR], 84; [CoL] ; [iNaturalist]
  • Spermophilus major Pallas, 1779;
  • S. m. major

    • rufescens
    • Citellus major argyropopuloi Bazhanov, 1947; ; TL: River Uil, NE of Caspian Sea

    • Spermophilus major major; [MPR], 84

• Spermophilus franklinii (Sabine, 1822) N.Great Plains, Alberta, Saskatchewan, Manitoba - Kansas, Illinois, Indiana. See [maps]

  • franklinii (Sabine, 1822); Trans. Linn. Soc. Lond. 13: 587; TL: Saskatschewan,Carlton House

  • Poliocitellus franklinii; [CoL] ; [iNaturalist]

• Spermophilus lateralis (Say, 1823) Rocyky Mountains (British Columbia - New Mexico), Columbia River - California, Nevada. See [maps]

  • lateralis (Say, 1823); in Long, Account of an Exped. from Pittsburgh to the Rocky Mtns., 2: 46; TL: Colorado, Fremont Co., Arkansas River, aboout 26 mi. below Canyon City

  • Callospermophilus lateralis; [CoL] ; [iNaturalist]
• Spermophilus madrensis (Merriam, 1901) SW.Chihuahua

  • madrensis (Merriam, 1901); Proc. Wash. Acad. Sci. 3: 563; TL: Mexico, Chihuahua, Sierra Madre, near Guadalupe y Calvo, 7000ft

  • Callospermophilus madrensis; [CoL] ; [iNaturalist]
• Spermophilus saturatus (Rhoads, 1895) W.Washington (Cascase Mts.), SW.British Columbia. See [maps]

  • saturatus (Rhoads, 1895); Proc. Acad. Nat. Sci. Philadelphia 47: 43; TL: Washington, Kittitas Co., Lake Kichelos (Keechelus), 8000ft

  • Callospermophilus saturatus; [CoL] ; [iNaturalist]

• Spermophilus mexicanus (Erxleben, 1777) S.New Mexico, W.Texas, Mexico. See [maps]

  • mexicanus (Erxleben, 1777); Syst. Regn. Anim.: 428; TL: Mexico, Toluca

  • Ictidomys mexicanus; [CoL] ; [iNaturalist]
• Spermophilus perotensis Merriam, 1893 Mexico (Veracruz, Puebla). See [maps]

  • Spermophilus perotensis Merriam, 1893; Proc. Biol. Soc. Washington, 8: 131; TL: Mexico, Veracruz, Perote
• Spermophilus spilosoma Bennett, 1833 C.Mexico - S.Texas, SW.South Dakota, NW.Arizona. See [maps]

  • Spermophilus spilosoma Bennett, 1833; Proc. Zool. Soc. Lond. 1833: 40; TL: Mexico, Durango, Durango City
• Spermophilus tridecemlineatus (Mitchill, 1821) Great Plains (C.Texas - E.Utah, Ohio - SC.Canada). See [maps] Thirteen-lined Ground Squirrel

  • tridecemlineatus (Mitchill, 1821); Med. Repos. (N.Y.) (n.s.) 6 (21): 248; TL: C.Minnesota

  • Ictidomys tridecemlineatus; [CoL] ; [iNaturalist]

• Spermophilus mohavensis Merriam, 1889 S.California (NW.Mohave Desert, Owens Valley). See [maps]

  • Spermophilus mohavensis Merriam, 1889; North Amer. Fauna 2: 15; TL: California, San Bernardino Co., near Rabbit Springs, about 15 mi E. Hesperia

  • Xerospermophilus mohavensis; [CoL] ; [iNaturalist]
• Spermophilus tereticaudus Baird, 1858 deserts (SE.California, S.Nevada, W.Arizona, NE.Baja California, Sonora). See [maps]

  • Spermophilus tereticaudus Baird, 1858; in Rep. Expl. Suvr. Railr. to Pacific, Mammals, 8 (1): 315; TL: California, Imperial Co., Old Fort Yuma

  • Xerospermophilus tereticaudus; [CoL] ; [iNaturalist]

20.2.2022 (36)

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.